Brass from another chamber even though it is supposed to be the same cartridge will often "NOT" fit another chamber for many reasons.
As far as the head space gauge, if the head space gauge is marked 22/250 AI, It will be .004 thousandths shorter than the standard 22/250 Head space gauge to hold the case during fire forming.
There are several different Shoulder angles on wildcatted cartridges and many are not 40 degrees
so it may have been chambered with one of them not a 40 degree reamer.
If you can please give us a larger picture of the case or take it to a competent Gun smith for him to solve you problems. It should fire a SAMMI 22/250 factory round without any problem.
PS: Never fire form brass at Max load data and Never, Never, with the bullet seated against the lands.
J E CUSTOM