I think what you are seeing us one of the most frustrating aspects of load development, and that is the collision of small sample size and the science of statistics.
Looking at the load with the best vertical...was the ES caused by just one shot or were the velocities evenly distributed across the range?
No offense but at 555 it is very possible for the shooter to account for a little of the vertical, and if that isn't frustrating enough, you can induce errors opposite the error caused by the velocity which can really throw you off on your analysis. Don't forget wind effects, I'm guessing it is pretty flat in Iowa but here in ky I have seen some vertical wind effects from wind flowing upslope and down slope and of course there is a small vertical wind effect caused by the horizontal component of wind.
Finally, it's in the nature of random dispersion that even with the same load you shoot five consecutive five shot groups in a row with your rifle that you will likely get five different group sizes, and if you measured the extreme spread of the vertical in those groups at 555 yards it is very possible to get an inch and half or difference between groups with all the real world sources of error.
This is a long winded way of saying that there is no definitive answer to your question and the only way to truly get truth data is to keep shooting and collecting the cleanest possible vertical spread data by reducing the other forms of error as much as possible. Low wind days, low mirage days, good clean trigger pulls and natural point of aim and good parallax adjustment, all that stuff you probably already know.
I think you should just try again with those things in mind, shoot as clean as you can, and see what you get.
The reason I know other sources of error are present is that even a 40fps velocity spread for a 180hybrid at 555 yards is only worth about 1.8 inches of vertical. Maybe on that last group you shot really well and didn't add much error to the inherent precision the rifle is capable of combined with the environmentals that day.
The good news is that it appears you have an accurate rifle and are shooting pretty well, so I think you will sort this out. If it were me, I'd not worry about the chronograph so much and shoot that last load again.
Also, are you shooting these groups with the magnetospeed in place? If so, shoot without it and use the magnetospeed to find the velocity after you sort out what the rifle likes.
Finally for what it is worth I reduced my ES slightly by graphiting the necks to get a more consistent bullet release.
Frustrating process at times but the fact is as shooters we rely on ES and SD from a single 5 shot group as Truth when it is mathematical fact with a sample size that small that the real long term SD of that load could be as much as 2.5 times off that number. The only way to know with confidence that you got a sample from the middle of the bell curve is to shoot more.